1. Ministry of Education Key Laboratory for Ecology of Tropical Islands, Key Laboratory of Tropical Animal and Plant Ecology of Hainan Province, College of Life Sciences Hainan Normal University Haikou China
2. Faculty of Agriculture, Forestry and Food Engineering Yibin University Yibin China
3. Institute of Ecology and Biological Resources Vietnam Academy of Science and Technology Hanoi Vietnam
4. Vietnam Academy of Science and Technology Graduate University of Science and Technology Hanoi Vietnam
5. State Key Laboratory of Genetic Resources and Evolution & Yunnan Key Laboratory of Biodiversity and Ecological Conservation of Gaoligong Mountain, Kunming Institute of Zoology Chinese Academy of Sciences Kunming China
6. Institute of Tropical Biology Vietnam Academy of Science and Technology Ho Chi Minh City Vietnam
7. Department of Herpetology California Academy of Sciences San Francisco California USA
8. Department of Herpetology American Museum of Natural History New York New York USA