1. Groningen Institute for Evolutionary Life Sciences; University of Groningen; P.O. Box 11103 Groningen 9700 CC The Netherlands
2. Faculty of Geosciences; Department of Earth Sciences; Utrecht University; Utrecht 3508 TA The Netherlands
3. Department of Coastal Systems; NIOZ Royal Netherlands Institute for Sea Research; Utrecht University; P.O. Box 59 Den Burg 1790 AB The Netherlands
4. Institute for Wetland and Water Research; Radboud University Nijmegen; Heyendaalseweg 135 Nijmegen 6525 AJ The Netherlands
5. Department of Marine Microbiology and Biogeochemistry; NIOZ Royal Netherlands Institute for Sea Research; Utrecht University; P.O. Box 59 Den Burg 1790 AB The Netherlands