1. Environmental Genomics Group; Max Planck Institute for Evolutionary Biology; 24 306 Plön Germany
2. Department of Evolutionary Theory; Max Planck Institute for Evolutionary Biology; 24 306 Plön Germany
3. Institute for Biology and Environmental Sciences; Carl von Ossietzky University; 26 111 Oldenburg Germany
4. Department of Genetics, Evolution, and Environment; University College London; London WC1E 6BT UK