1. National Engineering Laboratory for Industrial Enzymes and Tianjin Engineering Center for Biocatalytic Technology; Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences; Tianjin 300308 P. R. China
2. College of Bioengineering; Tianjin University of Science and Technology; Tianjin 300457 P. R. China
3. University of Chinese Academy of Sciences; Beijing 100049 P. R. China