1. Graduate School of Advanced Integrated Studies in Human Survivability; Kyoto University; Sakyo-ku Kyoto 606-8306 Japan
2. Div. of Applied Life Sciences; Graduate School of Agriculture, Kyoto University; Sakyo-ku Kyoto 606-8502 Japan
3. CREST, JST; Sakyo-ku Kyoto 606-8502 Japan
4. Japan Society for the Promotion of Science; Sakyo-ku Kyoto 606-8502 Japan