1. The Key Laboratory of Industrial Biotechnology, Ministry of Education; Jiangnan University; Wuxi 214122 People's Republic of China
2. National Engineering Laboratory for Cereal Fermentation Technology; Jiangnan University; Wuxi 214122 People's Republic of China
3. School of Biotechnology; Jiangnan University; Wuxi 214122 People's Republic of China
4. State Key Laboratory of Biological Fermentation Engineering of Beer; Tsingtao Brewery Co. Ltd; Qingdao 266100 People's Republic of China