1. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of BotanyChinese Academy of Sciences Kunming 650201 Yunnan P.R. China
2. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical GardenChinese Academy of Sciences Mengla 666303 Yunnan P.R. China
3. Department of Botany, National Museum of Natural HistorySmithsonian Institution Washington D.C. 20013‐7012 U.S.A.