1. Institute of Genetic Medicine; Newcastle University; Newcastle upon Tyne NE1 3BZ UK
2. Gavin Herbert Eye Institute; University of California Irvine; Irvine CA 92697 USA
3. Structural Biophysics Research Group; School of Optometry and Vision Sciences; Cardiff University; Cardiff CF24 4HQ Wales UK
4. Department of Ophthalmology; Royal Victoria Infirmary; Newcastle upon Tyne NE1 4LP UK