1. National Institute of Marine Sciences and Technologies, Salammbô Tunis Tunisia
2. Tunis El Manar University, Faculty of Sciences of Tunis, Research Laboratory of Diversity, Management and Conservation of Biological Systems, El Manar II Tunis Tunisia
3. University of Poitiers, Laboratory Ecology and Biology of Interactions (UMR CNRS 7267 EBI), Ecology Evolution Symbiosis Team Poitiers Cedex 9 France