1. Tunis El Manar University Higher Institute of Medical Technologies of Tunis Tunis Tunisia
2. Tunis El Manar University, Faculty of Sciences of Tunis, Research Laboratory of Diversity Management and Conservation of Biological Systems, El Manar II Tunis Tunisia
3. University of Poitiers, Laboratory Ecology and Biology of Interactions (UMR CNRS 7267 EBI) Team Ecology Evolution Symbiosis, University of Poitiers 5 street Albert Turpain, TSA 51106, 86073 Poitiers Cedex 9 Poitiers France