1. Department of Anatomy, School of Biomedical Sciences University of Otago Dunedin New Zealand
2. Cawthron Institute Nelson New Zealand
3. National Ecological Observatory Network Boulder Colorado USA
4. Department of Marine Science University of Otago Dunedin New Zealand
5. Department of Biochemistry University of Otago Dunedin New Zealand
6. Institute of Marine Science, University of Auckland Auckland New Zealand