1. Food Sciences, School of Biosciences; Sutton Bonington Campus, University of Nottingham LE12 5RD; UK
2. Barth Innovations Ltd, Hop Pocket Lane; Paddock Wood, Kent TN12 6DQ; UK
3. Brewing Science Department, School of Biosciences; Sutton Bonington Campus, University of Nottingham LE12 5RD; UK