1. Division of Biology Kansas State University 104 Ackert Hall Manhattan Kansas 66506 USA
2. U.S. Bureau of Reclamation Upper Colorado Regional Office 125 South State Street, Room 8100 Salt Lake City Utah 84138 USA
3. Department of Watershed Sciences Utah State University 5210 Old Main Hill Logan Utah 84322 USA
4. Department of Forest and Conservation Sciences University of British Columbia 3041‐2424 Main Mall Vancouver British Columbia V6T‐1Z4 Canada