1. Arizona Cooperative Fish and Wildlife Research Unit
2. University of ArizonaSchool of Natural Resources and the Environment ENR2 Rm N315, 1064 E. Lowell St. Tucson AZ 85721
3. U.S. Geological Survey, Arizona Cooperative Fish and Wildlife Research Unit
4. University of ArizonaSchool of Natural Resources and the Environment ENR2 Rm N315, 1064 E. Lowell St. Tucson AZ