1. Faculty of Health and Life Sciences; Department of Musculoskeletal Biology; Institute of Ageing and Chronic Disease; University of Liverpool; William Henry Duncan Building, 6 West Derby Street Liverpool L7 8TX United Kingdom
2. The MRC-Arthritis Research UK Centre for Integrated Research into Musculoskeletal Ageing (CIMA); Liverpool United Kingdom
3. Institute of Cellular Medicine; Newcastle University; Framlington Place Newcastle upon Tyne NE2 4HH United Kingdom