1. Key Laboratory for Plant Diversity and Biogeography of East Asia; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming Yunnan China
2. Plant Germplasm and Genomics Center; the Germplasm Bank of Wild Species; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming Yunnan China
3. Institute of Tibetan Plateau Research at Kunming; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming Yunnan China
4. College of Life Sciences; Henan Normal University; Xinxiang Henan China
5. College of Life Science; Yunnan Normal University; Kunming Yunnan China