1. Centre for Neuroregeneration; University of Edinburgh, Chancellor's Building, 49 Little France Crescent; Edinburgh EH16 4SB United Kingdom
2. University of Edinburgh, The Roslin Institute; Easter Bush Edinburgh EH25 9RG
3. University of Exeter Medical School, University of Exeter; Exeter UK
4. UK Dementia Research Institute, University of Edinburgh, Edinburgh Medical School, 47 Little France Crescent; Edinburgh, EH16 4TJ UK