1. Liverpool School of Tropical Medicine; Department of Clinical Sciences; Pembroke Place Liverpool UK L3 5QA
2. Independent consultant; Lawrenceville Georgia USA 30044
3. University of Technology Sydney; School of Life Sciences; Broadway NSW Australia
4. Macquarie University; Department of Biology; Ryde Australia
5. James Cook University; Australian Institute of Tropical Health and Medicine; Cairns Queensland Australia 4870