1. Faculty of Environment and Resource Studies; Mahidol University; Salaya Nakhon Pathom 73170 Thailand
2. Conservation Ecology Program; King Mongkut's University Thonbury; Bangkhuntien, Bangkok 10150 Thailand
3. Department of Biology; Faculty of Science, Mahidol University; Rachathewi Bangkok 10400 Thailand
4. Institute of Molecular Biosciences, Mahidol University; Salaya Nakhon Pathom 73170 Thailand
5. Ecology Laboratory; BIOTEC, National Science and Technology Development Agency; Khong 2 Pathum Thani 12123 Thailand