1. James Cook University Townsville QLD Australia
2. Deakin University Geelong VIC Australia
3. Institute of Marine and Antarctic Studies Hobart TAS Australia
4. School of Natural SciencesUniversity of Tasmania Hobart TAS Australia
5. CESIMAR (Centro Para el Estudio de Sistemas Marinos)–CENPAT–CONICET Puerto Madryn Chubut Argentina