1. Key Laboratory of Experimental Marine Biology Institute of Oceanology Chinese Academy of Sciences Nanhai Road 7 Qingdao 266071 P. R. China
2. Laboratory of Marine Biology and Biotechnology Qingdao National Laboratory for Marine Science and Technology Wenhai Road 1 Qingdao 266237 P. R. China
3. University of Chinese Academy of Sciences Yuquan Road 19 A Beijing 100049 P. R. China
4. Center for Ocean Mega-Science Chinese Academy of Sciences Nanhai Road 7 Qingdao 266071 P. R. China