1. School of Biological Engineering Dalian Polytechnic University Dalian People's Republic of China
2. Tianjin Institute of Industrial Biotechnology Chinese Academy of Sciences Tianjin China
3. Key Laboratory of Systems Microbial Biotechnology Chinese Academy of Sciences Tianjin People's Republic of China
4. School of Food Science and Technology, National Engineering Research Center of Seafood Dalian Polytechnic University Dalian People's Republic of China