1. Institute for Forest Resources & Environment of Guizhou Key Laboratory of Forest Cultivation in Plateau Mountain Area College of Forestry Guizhou University Guiyang 550025 China
2. Institute of Biological Research of Guizhou Guizhou Academy of Sciences Guiyang 550025 China
3. Guizhou Provincial Key Laboratory for Environment Guizhou Normal University Guiyang 550001 China