1. Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan
2. Chiba Prefectural Government Office, 1-1 Ichiba-cho, Chuo-ku, Chiba City, Chiba 260-8667, Japan
3. Forestry and Forest Products Research Institute, 1 Matsunosato, Tsukuba, Ibaraki 305-8687, Japan
4. Graduate School of Science, Kyoto University, Kitashirakawaoiwake-cho, Sakyo-ku, Kyoto-shi, Kyoto, 606-8502, Japan