1. Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, 106 Nanjing Road, Qingdao 266071, China
2. Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao National Laboratory for Marine Science and Technology, 1 Wenhai Road, Qingdao 266237, China
3. College of Fisheries and Life Science, Dalian Ocean University, 52 Heishijiao Road, Dalian 116023, China
4. Weifang Kenon Biological Technology Co., Ltd., 508 Zhujiang West Second Street, Weifang 262737, China