1. a Department of Biological Sciences and Centre for Environment and Society , University of Essex , Colchester, UK
2. b Unilever Research & Development, Colworth Laboratory , Sharnbrook, Bedford, UK
3. c Department of Soil Science , Rothamsted Research , Harpenden, Herts, UK
4. d ADAS , Gleadthorpe Grange, Meden Vale, Mansfield, Nottinghamshire, UK
5. e British Trust for Ornithology , Thetford, Norfolk, UK
6. f Unilever Rotterdam , Weena 455, Rotterdam, Netherlands