1. Nebraska Cooperative Fish and Wildlife Research Unit and School of Natural Resources, University of Nebraska-Lincoln, Nebraska 68583-0984, U.S.A.,
2. U.S. Geological Survey, Nebraska Cooperative Fish and Wildlife Research Unit and School of Natural Resources, University of Nebraska-Lincoln, Nebraska 68583-0984, U.S.A.
3. School of Natural Resources, University of Nebraska-Lincoln, Nebraska 68583-0984, U.S.A.