1. Centre for Mathematics and Physics in the Life Sciences and Experimental Biology, University College London, Gower Street, London, WC1E 6BT, United Kingdom
2. Department of Chemical Engineering, Centre for Process Systems Engineering, University College London, Torrington Place, London, WC1E 7JE, United Kingdom
3. Department of Neuroscience, Physiology & Pharmacology, University College London, Gower Street, London, WC1E 6BT, United Kingdom