1. Department of Biomedical Science University of Sheffield Sheffield UK
2. Neuroscience Institute University of Sheffield Sheffield UK
3. Oregon Hearing Research Center & Vollum Institute Oregon Health & Science University Portland OR USA
4. Present address: Department of Pathology Wexner Medical Center The Ohio State University Columbus OH USA
5. Present address: US Army Medical Department Activity‐Korea Camp Humphreys Republic of Korea
6. Mammalian Genetics Unit MRC Harwell Institute Harwell Campus Oxfordshire UK
7. Present address: UCL Ear Institute University College London London UK
8. Oregon Hearing Research Center Oregon Health & Science University Portland OR USA