1. Department of Genetics, Development and Cell Biology, Iowa State University, Ames, Iowa 50011
2. Interdepartmental Plant Biology, Iowa State University, Ames, Iowa 50011
3. Department of Crop and Soil Sciences, The University of Georgia, Athens, Georgia 30602
4. U.S. Department of Agriculture–Agriculture Research Service, Ithaca, New York 14853
5. Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York 14853