1. Department of Plant Biology, Michigan State University, East Lansing, Michigan 48824
2. Department of Horticulture, Virginia Tech, Blacksburg, Virginia 24061
3. Department of Plant, Soil, and Microbial Sciences, Michigan State University, East Lansing, Michigan 48824
4. Department of Horticulture, University of Wisconsin, Madison, Wisconsin 53706
5. Institute of Biotechnology, Provincial Key Laboratory of Agrobiology, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China