1. Institute of Radiation Breeding, National Institute of Agrobiological Sciences, Ohmiya-machi, Naka-gun, Ibaraki 319-2293, Japan
2. National Agricultural Research Center for the Western Region, National Agricultural Research Organization, Fukuyama, Hiroshima 721-8514, Japan
3. Kihara Institute for Biological Research, Yokohama City University, Maioka, Totsuka-ku, Yokohama 244-0813, Japan
4. Graduate School of Agricultural Science, Tohoku University, Sendai 981-8555, Japan