1. RIKEN Plant Science Center, Tsurumi, Yokohama 230-0045, Japan
2. Institut Jean-Pierre Bourgin, Unité Mixte de Recherche 1318, Institut National de la Recherche Agronomique-AgroParisTech, F-78026 Versailles cedex, France
3. Department of Disease and Stress Biology, John Innes Centre, Norwich NR4 7UH, United Kingdom