1. Department of Botany and Plant Sciences, University of California, Riverside, California 92521
2. Center for Plant Cell Biology, University of California, Riverside, California 92521
3. Department of Bioengineering, University of California, Riverside, California 92521
4. Department of Biological Sciences, Sung Kyun Kwan University, Jangan-gu, Suwon, Gyeonggi-do 440-746, Korea