1. Department of Botany, Graduate School of Science, Kyoto University, Kyoto 606-8502, Japan
2. Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma 630-0101, Japan
3. Kobe Advanced ICT Research Center, National Institute of Information and Communications Technology, Kobe 651-2492, Japan
4. Graduate School of Science, Osaka University, Toyonaka 560-0043, Japan
5. Graduate School of Frontier Biosciences, Osaka University, Suita 565-0871, Japan