1. Section of Horticulture, School of Integrative Plant Science, Cornell University, Ithaca, New York 14853
2. Beijing Advanced Innovation Center for Tree Breeding by Molecular Design, Beijing Forestry University, Beijing, 100083 China
3. Robert W. Holley Center for Agriculture and Health, United States Department of Agriculture, Agricultural Research Service, Cornell University, Ithaca, New York 14853
4. Department of Molecular Biology and Genetics, Cornell University, Ithaca, New York 14853
5. Department of Plant Sciences, University of California-Davis, Davis, California 95616