1. State Key Laboratory of Plant Genomics, National Center for Plant Gene Research, Institute of Genetics and Developmental Biology, Innovation Academy for Seed Design, Chinese Academy of Sciences, Beijing 100101, China
2. Chinese Academy of Sciences Center for Excellence in Biotic Interactions, University of Chinese Academy of Sciences, Beijing 100049, China
3. State Key Laboratory of Crop Biology, College of Agronomy, Shandong Agricultural University, Shandong Province, Tai’an 271018, China