1. Graduate School of Agricultural Science, Kyushu University, Higashi-ku, Fukuoka 812-8581, Japan
2. RIKEN Plant Science Center, Tsurumi-ku, Yokohama 230-0045, Japan
3. Faculty of Bioscience, Fukui Prefectural University, Matsuoka, Eiheiji-town, Fukui 910-1195, Japan
4. Graduate School of Agricultural Science, Tohoku University, Aoba-ku, Sendai 981-8555, Japan
5. Graduate School of Pharmaceutical Sciences, Chiba University, Chuo-ku, Chiba 260-8675, Japan
6. Department of Biochemistry and Molecular Biology, Michigan State University, East Lansing, Michigan 48824