1. Cranfield Soil and AgriFood Institute, School of Energy, Environment and Agrifood, Cranfield University, Cranfield, Bedfordshire, MK43 0AL, United Kingdom
2. Rothamsted Research, Harpenden, Hertfordshire, AL5 2JQ, United Kingdom
3. School of Life Sciences, University of Warwick, Gibbet Hill Campus, Coventry, CV4 7AL, United Kingdom