1. Biodiversity and Informatics, Landcare Research New Zealand, Auckland, 1072
2. Department of Mathematics and Statistics, University of Otago, Dunedin, 9054, New Zealand
3. Allan Wilson Centre for Molecular Ecology and Evolution, University of Otago, Dunedin, 9054, New Zealand
4. Centre for Reproduction and Genomics, Department of Anatomy, University of Otago, Dunedin, 9054, New Zealand