1. Division of Citrus Research, Institute of Fruit Tree and Tea Science, NARO, Okitsunakacho, Shimizu, Shizuoka 424-0292, Japan
2. The United Graduate School of Agricultural Science, Gifu University, 1-1 Yanagido, Gifu-shi, Gifu, 501-1193, Japan
3. Department of Bioresource Sciences, Faculty of Agriculture, Shizuoka University, 836 Ohya, Suruga, Shizuoka, 422-8529, Japan