1. State Key Laboratory of Grassland Agro‐ecosystems, Key Laboratory of Grassland Livestock Industry Innovation, Ministry of Agriculture and Rural Affairs, Engineering Research Center of Grassland Industry, Ministry of Education, College of Pastoral Agriculture Science and Technology Lanzhou University Lanzhou 730020 P. R. China
2. Hainan Key Laboratory for Sustainable Utilization of Tropical Bioresources, Institute of Tropical Agriculture and Forestry Hainan University Haikou 570228 P. R. China
3. International Research Centre for Environmental Membrane Biology Foshan University Foshan 528000 P. R. China
4. Tasmanian Institute of Agriculture University of Tasmania Private Bag 54 Hobart TAS 7001 Australia
5. School of Biological Sciences The University of Western Australia Perth WA 6009 Australia