1. Graduate School of Life and Environmental Sciences; University of Tsukuba; Tsukuba Ibaraki 305-8572 Japan
2. Institute for Molecular and Cellular Regulation; Gunma University; Maebashi Gunma 371-8512 Japan
3. Gunma University Initiative for Advanced Research; Gunma University; Maebashi Gunma 371-8511 Japan
4. Department of Cellular Biology; University of Georgia; Athens Georgia 30602 USA