1. Institute for Marine and Antarctic Studies; University of Tasmania; Private Bag 129 Hobart 7001 Tas. Australia
2. Australian Antarctic Division; 203 Channel Highway Kingston 7050 Tas. Australia
3. Australian Institute of Marine Science; The UWA Oceans Institute; 35 Stirling Highway Crawley WA 6009 Australia
4. School of Biological Sciences; University of Tasmania; Private Bag 5 Hobart 7001 Tas. Australia