1. Centre for Ecosystem Science School of Biological, Earth and Environmental Sciences UNSW Sydney NSW 2052 Australia
2. School of Mathematics and Statistics and Evolution & Ecology Research Centre The University of New South Wales Sydney NSW 2052 Australia
3. NSW National Parks and Wildlife Service Narrabri NSW 2390 Australia
4. NSW Office of Environment and Heritage Hurstville NSW 2220 Australia
5. Biological Sciences University of Southampton Southampton SO17 1BJ UK
6. School of Biosciences The University of Melbourne Melbourne Vic. 3010 Australia
7. Fenner School of Environment and Society The Australian National University Canberra ACT 2601 Australia
8. Department of Ecology, Evolution and Behavior University of Minnesota Saint Paul MN 55108 USA