1. Center for Applied Mathematics; Cornell University; Ithaca NY 14853 USA
2. Australian Research Council Centre of Excellence for Environmental Decisions; School of Biological Sciences; University of Queensland; St Lucia QLD 4072 Australia
3. Department of Entomology; Cornell University; Ithaca NY 14853 USA
4. Department of Ecology and Evolutionary Biology; Cornell University; Ithaca NY 14853 USA