1. Department of Life Sciences; Imperial College London, Silwood Park Campus; Buckhurst Road Ascot SL5 7PY UK
2. Department of Biological Chemistry and Crop Protection; Rothamsted Research; Harpenden Hertfordshire AL5 2JQ UK
3. College of Life and Environmental Science; University of Exeter, Penryn Campus; Penryn Cornwall TR10 9FE UK
4. Department of Organismal Biology; Queen Mary University of London; Mile End Road London E1 4NS UK