1. Department of Biology; University of Northern Iowa; Cedar Falls IA 50614-0294 USA
2. Department of Biology and Tallgrass Prairie Center; University of Northern Iowa; Cedar Falls IA 50614-0294 USA
3. National Laboratory for Agriculture and the Environment; USDA Agriculture Research Service; Ames IA 50011 USA
4. Department of Natural Resource Management; South Dakota State University; Brookings SD 57007-0001 USA