1. Institute for Conservation Research; 15600 San Pasqual Valley Road Escondido CA USA
2. U.S. Geological Survey; Western Ecological Research Center; 160 North Stephanie Street Henderson NV USA
3. U.S. Fish and Wildlife Service; Desert Tortoise Recovery Office; 1340 Financial Boulevard #234 Reno NV USA
4. Department of Geography; University of Nevada Reno; 1664 N. Virginia Street Reno NV USA