1. Department of Biological Sciences; Gombe State University; Tudan Wada Street Gombe Gombe State Nigeria
2. School of Biological Sciences; The University of Queensland; Brisbane Qld 4072 Australia
3. School of Mathematics and Statistics; University of Canterbury; PB 4800 Christchurch New Zealand
4. UMR 7179 MECADEV CNRS-MNHN; Département Adaptations du vivant; Muséum National d'Histoire Naturelle; 1 avenue du Petit Château 91800 Brunoy France
5. School of Biological Sciences; University of Canterbury; PB 4800 Christchurch New Zealand
6. Nigerian Montane Forest Project; Yelwa Village Taraba State Nigeria